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Ecologist Hiroshi Saito has argued that the restoration of oyster reefs can significantly improve conditions in coastal estuaries. Oysters are natural filter feeders, capable of processing gallons of water daily and removing excess nitrogen and suspended sediments. ______ the reefs themselves provide a structural habitat that shelters juvenile fish, crabs, and other marine species. Which choice completes the text with the most logical transition?

  1. Thus,
  2. Specifically,
  3. Conversely,
  4. Additionally,

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