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Environmental scientists monitoring the health of coral reefs have noted that certain species of algae are increasingly dominating degraded reef systems. While these algae provide some structural cover, they do not offer the complex niches required by diverse fish populations. ______ the proliferation of these algae is often a sign of a reef in transition toward a less diverse ecosystem.

  1. Alternatively,
  2. Regardless,
  3. In short,
  4. Indeed,

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