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The 1977 discovery of hydrothermal ______ long-standing assumptions about where life on Earth can survive. Which choice completes the text so that it conforms to the conventions of Standard English?

  1. vents, openings on the seafloor that gush superheated water, upended
  2. vents, openings on the seafloor that gush superheated water — upended
  3. vents — openings on the seafloor that gush superheated water, upended
  4. vents openings on the seafloor that gush superheated water, upended

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