medium · SAT Reading & Writing

For many years, it was assumed that the deep ocean was a 'biological desert' because the lack of sunlight made photosynthesis impossible. _______, the 1977 discovery of hydrothermal vents revealed entire ecosystems thriving on chemical energy from the Earth's interior, independent of the sun's rays.

Which choice completes the text with the most logical transition?

  1. Nevertheless
  2. Therefore
  3. Indeed
  4. Specifically

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