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For decades, biologists believed that the development of complex multicellular life required a significant and sudden increase in atmospheric oxygen. ________ recent analysis of ancient rock layers has revealed that multicellular organisms may have appeared long before the primary rise in oxygen levels, suggesting that other factors may have triggered the transition.

Which choice completes the text with the most logical transition?

  1. Furthermore,
  2. In contrast,
  3. Specifically,
  4. Accordingly,

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